{
  "contentStorage": "postgres",
  "title": "卡罗尔·格雷德 · Beziehungen",
  "description": "卡罗尔·格雷德是与布莱克本共同发现端粒酶的分子生物学家，2009年诺贝尔生理学或医学奖得主之一。",
  "url": "/de/people/carol-w-greider/relation.html",
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    "type": "person",
    "title": "卡罗尔·格雷德",
    "slug": "carol-w-greider",
    "aliases": [
      "Carol W. Greider"
    ],
    "summary": "卡罗尔·格雷德是与布莱克本共同发现端粒酶的分子生物学家，2009年诺贝尔生理学或医学奖得主之一。",
    "status": "published",
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        "id": "s1",
        "title": "Carol W. Greider – Facts",
        "author": "The Nobel Foundation",
        "url": "https://www.nobelprize.org/prizes/medicine/2009/greider/facts/"
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      {
        "id": "s2",
        "title": "The Nobel Prize in Physiology or Medicine 2009",
        "author": "The Nobel Foundation",
        "url": "https://www.nobelprize.org/prizes/medicine/2009/summary/"
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        "id": "s3",
        "title": "卡罗尔·格雷德 – Biographical",
        "author": "The Nobel Foundation",
        "url": "https://www.nobelprize.org/prizes/medicine/2009/greider/biographical/"
      },
      {
        "id": "s4",
        "title": "卡罗尔·格雷德",
        "author": "Wikipedia",
        "url": "https://zh.wikipedia.org/wiki/%E5%8D%A1%E7%BD%97%E5%B0%94%C2%B7%E6%A0%BC%E9%9B%B7%E5%BE%B7"
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    "createdAt": "2026-11-01T03:38:01+08:00",
    "updatedAt": "2026-11-02T14:48:34+08:00",
    "updatedBy": "user:admin",
    "revision": 4,
    "body": "**卡罗尔·格雷德**（Carol W. Greider，1961年4月15日—）出生于圣迭戈（美国）。\n\n## 生平与职业生涯\n\n格雷德在加州大学圣迭戈分校获博士学位，在冷泉港实验室从事博士后研究，后任约翰斯·霍普金斯大学教授。[^s3][^s4]\n\n## 主要成就\n\n格雷德与布莱克本共同发现端粒酶，揭示了染色体末端端粒的延长机制，为理解细胞衰老与癌症发生提供了关键线索。[^s3][^s4]\n\n## 获奖\n\n2009 年诺贝尔生理学或医学奖由 3 位得主共享，卡罗尔·格雷德占其中 1/3 份，官方获奖理由（官方原文）：“for the discovery of how chromosomes are protected by telomeres and the enzyme telomerase”[^s1]。获奖时所属机构为 Johns Hopkins University School of Medicine，所在地为巴尔的摩（美国）[^s1]。\n\n## 参见\n\n[[work_nobel_prize_2009|2009 年诺贝尔奖]]、[[concept_nobel_prize|诺贝尔奖]]",
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    "birth": {
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      "month": 4,
      "day": 15,
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      "display": "1961年4月15日"
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    "occupations": [
      "分子生物学家"
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      "label": "Birth",
      "value": "1961年4月15日"
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